The Eastern Transverse Ranges, adjacent to and southeast of the big left bend of the San Andreas fault, southern California, form a crustal block that has rotated clockwise in response to dextral shear within the San Andreas system. Previous studies have indicated a discrepancy between the measured magnitudes of left slip on through-going east-striking fault zones of the Eastern Transverse Ranges and those predicted by simple geometric models using paleomagnetically determined clockwise rotations of basalts distributed along the faults. To assess the magnitude and source of this discrepancy, we apply new gravity and magnetic data in combination with geologic data to better constrain cumulative fault offsets and to define basin structure for the block between the Pinto Mountain and Chiriaco fault zones. Estimates of offset from using the length of pull-apart basins developed within left-stepping strands of the sinistral faults are consistent with those derived by matching offset magnetic anomalies and bedrock patterns, indicating a cumulative offset of at most ???40 km. The upper limit of displacements constrained by the geophysical and geologic data overlaps with the lower limit of those predicted at the 95% confidence level by models of conservative slip located on margins of rigid rotating blocks and the clockwise rotation of the paleomagnetic vectors. Any discrepancy is likely resolved by internal deformation within the blocks, such as intense deformation adjacent to the San Andreas fault (that can account for the absence of basins there as predicted by rigid-block models) and linkage via subsidiary faults between the main faults. ?? 2009 Geological Society of America.