Knopf et al. found that summer cattle grazing has an adverse effect on the presence of certain willow-dependent songbirds. Pastures that have historical summer grazing no longer have the Willow flycatcher, Lincoln's sparrow and the White-crowned sparrow present. Yet in these same areas, birds like the American Robin, Brown-headed cowbird and the Red-winged blackbird have increased in density. One possible answer for the decrease in some songbirds is the fact that the main focus of the Arapaho National Wildlife Refuge is on waterfowl habitat, which requires large amounts of open space (opposite of desirable songbird habitat).